BIOLOGIA MOLECOLARE DELL'EMOSTASI


Prodotti della ricerca

 

Modulation of factor V levels in plasma by polymorphisms in the C2 domain

Autori: Scanavini D, Girelli D, Lunghi B, Martinelli N, Legnani C, Pinotti M, Palareti G, Bernardi F.
Rivista: Arterioscler Thromb Vasc Biol. Volume: 24 Anno: 2004

Abstract: OBJECTIVE: Functional polymorphisms contributing to coagulation factor levels are preferential markers for association studies aimed at identifying prothrombic genetic components. METHODS AND RESULTS: Factor V (FV) microsatellite genotypes were found to be associated with FV levels (P=0.003). Single nucleotide polymorphisms analysis and sequencing of the promoter and of coding regions identified two polymorphisms (Met2120Thr, Asp2194Gly) present in 20% of the population (n=1013) that are responsible for genotype-phenotype associations. The effect of the Met2120Thr polymorphism, both in plasma (mean reduction of FV level in the heterozygous condition: 25%) and in recombinant FV studies (34% reduction), was comparable to that of the Asp2194Gly change (20% and 34%, respectively). The study of 10 subjects with a rare genotype indicated that the Asp2194Gly substitution is the functional determinant of the reduced FV levels associated with the FVHR2 haplotype. Among Leiden carriers, the doubly heterozygous condition for FV2120Thr was found to be associated with a significantly increased activated protein-C resistance (APCR) (P<0.05), and the doubly heterozygous condition for FV2194Gly was found to be more frequent (P=0.009) in symptomatic than in asymptomatic subjects. CONCLUSIONS: Extensive analysis of FV polymorphisms indicated that changes in the C2 domain modulate FV levels and might increase APCR and thrombotic risk in FV Leiden carriers through a pseudohomozygous mechanism.

Breve commento esplicativo: L’analisi di diversi polimorfismi localizzati nel dominio C2 del fattore V (FV) della coagulazione ha mostrato che questi possono modulare i livelli plasmatici del FV stesso ed incrementare la resistensa alla proteina C attivata. La conseguenza è l’aumento del rischio di trombosi nei soggeti già portatori della mutazione Leiden. Questi polimorfismi funzionali possono essere utilizzati come marcatori genetici di rischio trombotico.



Coinheritance of Factor V (FV) Leiden enhances thrombin formation and is associated with a mild bleeding phenotype in patients homozygous for the FVII 9726+5G>A (FVII Lazio) mutation

Autori: Castoldi E, Govers-Riemslag JW, Pinotti M, Bindini D, Tans G, Berrettini M, Mazzucconi MG, Bernardi F, Rosing J.
Rivista: Blood Volume: 102 Anno: 2003

Abstract: We investigated the role of thrombophilic mutations as possible modifiers of the clinical phenotype in severe factor VII (FVII) deficiency. Among 7 patients homozygous for a cross-reacting material-negative (CRM-) FVII defect (9726+5G>A, FVII Lazio), the only asymptomatic individual carried FV Leiden. Differential modulation of FVII levels by intragenic polymorphisms was excluded by a FVII to factor X (FX) gene haplotype analysis. The coagulation efficiency in the FV Leiden carrier and a noncarrier was evaluated by measuring FXa, FVa, and thrombin generation after extrinsic activation of plasma in the absence and presence of activated protein C (APC). In both patients coagulation factor activation was much slower and resulted in significantly lower amounts of FXa and thrombin than in a normal control. However, more FXa and thrombin were formed in the plasma of the patient carrying FV Leiden than in the noncarrier, especially in the presence of APC. These results were confirmed in FV-FVII doubly deficient plasma reconstituted with purified normal FV or FV Leiden. The difference in thrombin generation between plasmas reconstituted with normal FV or FV Leiden gradually decreased at increasing FVII concentration. We conclude that coinheritance of FV Leiden increases thrombin formation and can improve the clinical phenotype in patients with severe FVII deficiency.

Breve commento esplicativo: In questo studio si è mostrato come la presenza del fenotipo Leiden aumenta la produzione di trombina migliorando il quadro clinico di pazienti aventi difetto di fattore VII (FVII). Ciò evidenzia come una mutazione che normalmente aumenta il rischio di trombosi possa, in alcuni casi, compensare la carenza di un fattore procoagulante.



Reduced activation of the Gla19Ala FX variant via the extrinsic coagulation pathway results in symptomatic CRMred FX deficiency

Autori: Pinotti M, Marchetti G, Baroni M, Cinotti F, Morfini M, Bernardi F.
Rivista: Thrombosis and haemostasis Volume: 88 Anno: 2002

Abstract: We characterized a symptomatic CRMred factor X (FX) deficiency produced by the Glu19Ala mutation in the gamma-carboxyglutamic-rich domain. FX activity levels in plasma were markedly reduced in prothrombin time assays (< 1-5%), whereas in activated partial thromboplastin assays (16%) and in RVV assays (17%) the reduction in activity mirrored that in antigen levels (17%). Activation of recombinant 19Ala-FX by factor IXa/factor VIIIa or RVV, and the activity in thrombin generation assays, were comparable to those of wild-type FX. Differently, complete activation of recombinant 19Ala-FX required a factor VIIa/TF concentration 30-fold higher than that of wild-type FX. The recombinant FVIIa significantly reduced PT values in 19Ala-FX reconstituted plasma, thus suggesting an alternative approach for treatment of FX deficiencies characterized by defective FX activation. The study of this FX deficiency provides an "in vivo" and "in vitro" model for the investigation of Gla domain interactions.

Breve commento esplicativo: Sono stati caratterizzati pazienti con difetto di fattore X (FX) della coagulazione dovuto ad una mutazione che ne coinvolge il dominio ricco in gamma-carbossiglutammico (Gla). Il dato più ecclatante di questo studio riguarda il fatto che la completa attivazione del FX mutato richiede una quantità di fattore VIIa/fattore tissutale 30 volte superiore alla forma normale, mentre l’attivazione tramite fattore IXa/fattore VIIIa è confrontabile tra le due diverse forme. Ciò può essere un valido suggerimento per studiare le interazioni tra il FX e gli altri fattori della coagulazione che coinvolgono il dominio Gla.


Combinations of 4 mutations (FV R506Q, FV H1299R, FV Y1702C, PT 20210G/A) affecting the prothrombinase complex in a thrombophilic family

Autori: Castoldi E, Simioni P, Kalafatis M, Lunghi B, Tormene D, Girelli D, Girolami A, Bernardi F.
Rivista: Blood Volume: 96 Anno: 2000

Abstract: The study of the molecular bases of thrombophilia in a large family with 4 symptomatic members is reported. Three thrombophilic genetic components (FV R506Q, FV H1299R, and PT 20210G/A), all affecting the activity of the prothrombinase complex, were detected alone and in combination in various family members. In addition, a newly identified missense mutation (factor V [FV] Y1702C), causing FV deficiency, was also present in the family and appeared to enhance activated protein C (APC) resistance in carriers of FV R506Q or FV H1299R by abolishing the expression of the counterpart FV allele. The relationships between complex genotypes, coagulation laboratory findings, and clinical phenotypes were analyzed in the family. All symptomatic family members were carriers of combined defects and showed APC resistance and elevated F1 + 2 values. Evidence for the causative role of the FV Y1702C mutation, which affects a residue absolutely conserved in all 3 A domains of FV, factor VIII, and ceruloplasmin, relies on (1) the absolute cosegregation between the mutation and FV deficiency, both in the family and in the general population; (2) FV antigen and immunoblot studies indicating the absence of Y1702C FV molecules in plasma of carriers of the mutation, despite normal levels of the FV Y1702C messenger RNA; and (3) molecular modeling data that support a crucial role of the mutated residue in the A domain structure. These findings help to interpret the variable penetrance of thrombosis in thrombophilic families and to define the molecular bases of FV deficiency.

Breve commento esplicativo: In questo sono state identificate mutazioni che determinano cali di attività del complesso protrombinasico o riduzione dei livelli di fattore V (FV). Quando queste compaiono in associazione ad altre mutazioni del FV possono determinare una maggiore resistenza alla proteina C attivata. Ciò suggerisce di estendere lo screenig a tutte queste varianti per avere un quadro più dettagliato delle possibili cause genetiche della patologia.



Daily and Circadian Rhythms of Tissue Factor Pathway Inhibitor and Factor VII Activity

Autori: Pinotti M, Bertolucci C, Portaluppi F, Colognesi I, Frigato E, Foa A, Bernardi F.
Rivista: Arterioscler Thromb Vasc Biol. Volume: 16 Anno: 2004

Abstract: OBJECTIVE: Diurnal variations in levels of factor VII (FVII), FVIII, proteins C and S, antithrombin, plasminogen activator inhibitor-1, prothrombin fragment F1+2, and D-dimers in healthy humans point to the existence of circadian rhythms of coagulation factors. We sought for temporal fluctuations of tissue factor pathway inhibitor (TFPI) activity in human and mouse plasma. METHODS AND RESULTS: TFPI activity showed significant daily variations with highest levels in the morning in healthy men (+11%) and in mice at the light-to-dark transition (+63%), the beginning of the physically active period. Variations in FVII activity paralleled those in TFPI. In mice, the feeding schedule had a strong impact on these rhythms. Although restricted feeding and fasting shifted the peak of TFPI, the FVII peak disappeared. Investigation of temporal fluctuations in constant darkness indicated the existence of daily rhythms for TFPI and of true circadian rhythms for FVII. CONCLUSIONS: For the first time, we report, both in humans and mice, temporal variations in TFPI activity. The coherent variations in FVII and TFPI activity could interplay to maintain the coagulation equilibrium. The chronobiological patterns should be considered to analyze activity levels of these factors. Moreover, the mouse model could be exploited to investigate modifiers of coagulation rhythms potentially associated to morning peaks of cardiovascular events.

Breve commento esplicativo: In questo studio abbiamo mostrato la variazione secondo ritmi circadiani dell’inibitore della via del fattore tissutale (TFPI) e del fattore VII (FVII) della coagulazione, sia in modello murino che nell’uomo. Il passo successivo potrebbe riguardare l’analisi di altri fattori coinvolti nel processo emostatico per comprendere la relazione che hanno con il picco di eventi cardiovascolari che si verifica preferenzialmente al mattino.

 

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